Himalayan uplift and the strengthening monsoon (Miocene)
The collision of India with Eurasia pushed up the Himalayas and reshaped the river systems draining them. Around 10 million years ago the South Asian monsoon intensified, creating wet and dry seasons across terrain that already ranged from tropical valley to alpine slope.
New ecological niches appeared along elevation gradients and between river valleys.
The paper dates Jingophrys's rapid species splitting to roughly 11.3 to 4.1 million years ago, overlapping this period.
It explains why so many similar frogs can live within a few kilometres of each other: the mountains and rivers kept dividing populations.
